Ben Creisler | 11 Feb 01:26 2016

Dinosaur Footprints in Zizhou, Shaanxi Province, China + Jurassic angiosperm from China + more

Ben Creisler
bcreisler <at>

A number of recent papers. (Downloads for the Chinese papers are slow...:):

WANG Baopeng, LI Jianjun, BAI Zhiqiang, GAO Junmin, DONG Shurong, HU
Bailin, ZHAO Suiqin and CHANG Jieqiong (2016),
Research on Dinosaur Footprints in Zizhou, Shaanxi Province, China.
Acta Geologica Sinica 90(1):1-18
http: //

[pdf is free]

In recent years, the discoveries of dinosaur footprints have been
successively reported from Dianshi Town, Zizhou City, Shaanxi
Province. The footprints include the tracks of theropod, ornithopod
and sauropod as well. Actually, the dinosaur footprints were found by
local inhabitants much earlier in history, but not for science. The
slabs bearing the dinosaur footprints were collected for domestic use,
such as building stones, millstones, cellar covers, sheepfold fences,
windlass holders, etc. This paper is to describe the dinosaur
footprints on both sides of three slabs used for cellar covers,
sheepfold fences and windlass holders by the local people. 24 dinosaur
footprints and 4 trackways have been recognized and all of them belong
to theropod. Four kinds of dinosaur footprints are identified,
including 1 new ichnogenus and 2 new ichnospecies: (1) Shanbeipus
caudatus ichnogen. et ichnosp. nov.; (2) Pengxianpus yulinensis
ichnosp. nov.; (3) Eubrontes ichnosp; (4) Kayentapus ichnosp.
Dinosaur-footprint-bearing beds were initially identified as the
Fuxian Formation of the Lower Jurassic. Multipal dinosaur footprints
(Continue reading)

Richard W. Travsky | 11 Feb 00:56 2016

Jurassic butterflies

Don't think I've seen this mentioned; it's pretty

The evolutionary convergence of mid-Mesozoic lacewings and Cenozoic 


Mid-Mesozoic kalligrammatid lacewings (Neuroptera) entered the fossil 
record 165 million years ago (Ma) and disappeared 45 Ma later. Extant 
papilionoid butterflies (Lepidoptera) probably originated 80–70 Ma, long 
after kalligrammatids became extinct. Although poor preservation of 
kalligrammatid fossils previously prevented their detailed morphological 
and ecological characterization, we examine new, well-preserved, 
kalligrammatid fossils from Middle Jurassic and Early Cretaceous sites 
in northeastern China to unravel a surprising array of similar 
morphological and ecological features in these two, unrelated clades. We 
used polarized light and epifluorescence photography, SEM imaging, 
energy dispersive spectrometry and time-of-flight secondary ion mass 
spectrometry to examine kalligrammatid fossils and their environment. We 
mapped the evolution of specific traits onto a kalligrammatid phylogeny 
and discovered that these extinct lacewings convergently evolved wing 
eyespots that possibly contained melanin, and wing scales, elongate 
tubular proboscides, similar feeding styles, and seed–plant 
associations, similar to butterflies. Long-proboscid kalligrammatid 
lacewings lived in ecosystems with gymnosperm–insect relationships and 
likely accessed bennettitalean pollination drops and pollen. This system 
(Continue reading)

Paul P | 10 Feb 22:14 2016

Re: A New Ankylosaurid Dinosaur from the Upper Cretaceous (Kirtlandian) of New Mexico with Implications for Ankylosaurid Diversity in

Er, what?  Are you just reposting a very old link, or was there 
something else in that error-message block down below..?

On Wed, 2/10/16, Ben Creisler <bcreisler <at>> wrote:

 Subject: Re: A New Ankylosaurid Dinosaur from the Upper Cretaceous (Kirtlandian) of New Mexico with
Implications for Ankylosaurid Diversity in
 To: "mjmbego" <mjmbego <at>>, dinosaur <at>
 Date: Wednesday, February 10, 2016, 9:44 AM

 Ben Creisler
 bcreisler <at>

 Thanks for
 the mention. In fact, Ziapelta was posted on the DML back in

 On Wed, Feb 10, 2016 at
 2:17 AM, mjmbego . <mjmbego <at>>
> * * * * * * * * * * * * * * * * * * * * * * * * * * * * * *
> *         ---REMAINDER OF MESSAGE TRUNCATED---            *
> *     This post contains a forbidden message format       *
> *  (such as an attached file, a v-card, HTML formatting)  *
> *    This Mail List at USC.EDU only accepts PLAIN TEXT    *
> * If your postings display this message your mail program *
(Continue reading)

Ben Creisler | 10 Feb 21:52 2016

Viavenator, new abelisaurid theropod from Upper Cretaceous of Argentina

Ben Creisler
bcreisler <at>

A new paper.
Thanks to Rubén Darío Juárez Valieri for bringing this paper to my attention.

Leonardo S. Filippi, Ariel H. Méndez, Rubén D. Juárez Valieri &
Alberto C. Garrido (2016)
A new brachyrostran with hypertrophied axial structures reveals an
unexpected radiation of latest Cretaceous abelisaurids.
Cretaceous Research 61: 209–219
http: //

A well preserved skeleton of a new abelisaurid is reported here. The
holotype of Viavenator exxoni was found in the outcrops of the Bajo de
la Carpa Formation (Santonian, Upper Cretaceous), northwestern
Patagonia, Argentina. This new taxon belongs to the South American
clade of abelisaurids, the brachyrostrans. The current phylogenetic
analysis places it as the basalmost member of a new clade of derived
brachyrostrans, named Furileusauria, characterized by distinctive
cranial, axial and appendicular anatomical features. The Santonian age
of Viavenator allows filling the stratigraphic gap exhibited between
the basal brachyrostrans of Cenomanian–Turonian age, and the derived
forms from the Campanian-Maastrichtian. The evolution of abelisaurids
during the Late Cretaceous, faunal replacements, and the adaptive
radiation that occurred during that period of time in South America
are discussed.

(Continue reading)

Ben Creisler | 10 Feb 20:08 2016

Skull Sutures Are Ambiguous Maturity Indicators for Dinosauria

Ben Creisler
bcreisler <at>

New in PLoS ONE:

Alida M. Bailleul, John B. Scannella, John R. Horner& David C. Evans (2016)
Fusion Patterns in the Skulls of Modern Archosaurs Reveal That Sutures
Are Ambiguous Maturity Indicators for the Dinosauria.
PLoS ONE 11(2): e0147687.
http: //

The sutures of the skulls of vertebrates are generally open early in
life and slowly close as maturity is attained. The assumption that all
vertebrates follow this pattern of progressive sutural closure has
been used to assess maturity in the fossil remains of non-avian
dinosaurs. Here, we test this assumption in two members of the Extant
Phylogenetic Bracket of the Dinosauria, the emu, Dromaius
novaehollandiae and the American alligator, Alligator
mississippiensis, by investigating the sequence and timing of sutural
fusion in their skulls. As expected, almost all the sutures in the emu
skull progressively close (i.e., they get narrower) and then
obliterate during ontogeny. However, in the American alligator, only
two sutures out of 36 obliterate completely and they do so during
embryonic development. Surprisingly, as maturity progresses, many
sutures of alligators become wider in large individuals compared to
younger, smaller individuals. Histological and histomorphometric
analyses on two sutures and one synchondrosis in an ontogenetic series
of American alligator confirmed our morphological observations. This
pattern of sutural widening might reflect feeding biomechanics and
(Continue reading)

mjmbego . | 10 Feb 11:17 2016
Ben Creisler | 10 Feb 03:10 2016

Theropods from Idaho + Joe the baby Parasaurolophus at Dino Expo in Japan + more

Ben Creisler
bcreisler <at>

Some recent items (urls are separated):

Theropods from Idaho

http: //


Dracoraptor, with video

http: //


Joe, the baby Parasaurolophus from Utah, to be part of international
dinosaur exhibit at National Museum of Nature and Science in Tokyo,

http: //


Scotty the Tyrannosaurus is on display, as well as Spinosaurus

Dinosaur Expo link (in Japanese)

http: //
(Continue reading)

Ben Creisler | 9 Feb 00:09 2016

Pterosaur talk video + Pteros, new online pterosaur encyclopedia + more

Ben Creisler
bcreisler <at>

Some recent items. The urls are separated to prevent rewriting and redirection.

Pterosaur talk video for Royal Tyrrell Museum of Palaeontology speaker series

Donald Henderson -
"Over the Heads of Dinosaurs-Pterosaurs"

https: //


Pterosaurs, and Pteros, new online pterosaur encyclopedia

http: //


Pedro Mocho is in Teruel studying Turiasaurus for a doctoral
dissertation "Evolutionary history of the Upper Jurassic sauropods of
the Lusitanian Basin of Portugal, "and will announce and describe a
new sauropod soon (in Spanish)

http: //


More on Notocolossus (in Spanish)
(Continue reading)

Ben Creisler | 8 Feb 18:18 2016

Didactyl theropod tracks from Cretaceous Dinosaur Ridge in Colorado

Ben Creisler
bcreisler <at>

A new paper:
(url is separated)

Martin G. Lockley, Lida Xing, Neffra A. Matthews & Brent H. Breithaupt (2016)
Didactyl raptor tracks from the Cretaceous, Plainview Sandstone at
Dinosaur Ridge.
Cretaceous Research 61: 161–168
http: //

Two natural casts of two-toed (didactyl) tracks from the Cretaceous
(Albian) Plainview Sandstone (Plainview Member) of the South Platte
Formation (Dakota Group) at Dinosaur Ridge, Colorado are attributed to
deinonychosaurian theropod dinosaurs and placed in the ichnogenus
Dromaeosauripus. This is both the first report of tracks from this
unit in the Dinosaur Ridge area and the first report of
deinonychosaurian tracks from Colorado. It is also only the third
report of this track type from North America. The rarity of tracks
from the Albian-aged, Plainview Sandstone (Dakota Group Sequence 2)
contrasts with their abundance in the upper (Cenomanian) part of the
overlying South Platte Formation (Dakota Group Sequence 3), which has
yielded more than 120 sites mostly in Colorado, giving rise to the
“Dinosaur Freeway” concept. As no deinonychosaurid tracks are known
from the sequence 3 part of the South Platte Formation, despite the
large vertebrate and invertebrate ichnological database available, it
is evident that the sparse vertebrate ichnofauna from the Plainview
Member (Sequence 2) is inherently different. This striking difference
(Continue reading)

Paul P | 8 Feb 00:25 2016

Re: Skull ornamentation in juvenile Pachycephalosaurus fossils from Upper Cretaceous Hell Creek Formation

That link doesn't work [either].

On Sun, 2/7/16, Ben Creisler <bcreisler <at>> wrote:

 Subject: Re: Skull ornamentation in juvenile Pachycephalosaurus fossils from Upper Cretaceous Hell
Creek Formation
 To: dinosaur <at>
 Date: Sunday, February 7, 2016, 12:31 AM

 Sorry. Same fix to this
 ref to prevent the url rewrite and redirect:

 Mark B. Goodwin & David C.
 Evans (2016)
 The early expression of
 squamosal horns and parietal ornamentation
 confirmed by new end-stage juvenile
 Pachycephalosaurus fossils from
 the Upper
 Cretaceous Hell Creek Formation, Montana.
 Journal of Vertebrate Paleontology (advance
 online publication)
 http: //

 On Sat, Feb 6, 2016 at 10:27
 PM, Ben Creisler <bcreisler <at>>
(Continue reading)

Ben Creisler | 7 Feb 07:37 2016

Brachyprosopus (dicynodont from Middle Permian of South Africa) redescribed

Ben Creisler
bcreisler <at>

A new non-dino paper:

Kenneth D. Angielczyk, Bruce S. Rubidge, Michael O. Day & Florence Lin (2016)
A reevaluation of Brachyprosopus broomi and Chelydontops altidentalis,
dicynodonts (Therapsida, Anomodontia) from the middle Permian
Tapinocephalus Assemblage Zone of the Karoo Basin, South Africa.
Journal of Vertebrate Paleontology (advance online publication)
http: //

Brachyprosopus broomi was described in 1937 based on a specimen from
the Tapinocephalus Assemblage Zone (Karoo Basin, South Africa), but it
was largely overlooked by subsequent workers. We have identified
several new specimens that show that Brachyprosopus is a valid taxon.
An autapomorphy for the taxon is a curled lateral edge of the
squamosal that forms a lateral wall of the external adductor fossa.
Other important characters are absence of anterior median palatal
ridges; maxillary tooth rows bounded laterally by a shelf; unfused
vomers; raised margins of the interpterygoid vacuity; broad
intertemporal region; pineal boss; dentary tables; and a long, wide
posterior dentary sulcus that extends posterior to the dentary teeth.
Chelydontops altidentalis is a junior synonym of B. broomi. A
phylogenetic analysis places Brachyprosopus among basal dicynodonts,
not as a close relative of Endothiodon. It is noteworthy that some
characters, such as well-developed medial maxillary tooth rows and the
shape of the palatines, are shared by Brachyprosopus, Pristerodon,
Endothiodon, and Niassodon, hinting that a final resolution of
(Continue reading)